0001193125-22-276921 | 10-Q | XBRL Viewer

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Share Based Compensation- Summary Of Share Based Compensation Performance Shares Award Outstanding Activity (Detail) - Performance Stock Units [Member]	9 Months Ended
	Sep. 30, 2022 shares
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Outstanding as of December 31, 2021:	745,000
Issued	530,000
Forfeited/canceled	(145,000)
Exercised	0
Outstanding as of September 30, 2022:	1,130,000

Leases, Commitments, and Contingencies - Additional Information (Detail) $ in Millions	Sep. 30, 2022 USD ($) ft²
Roseville, MN Lease [Member]
Other Commitments [Line Items]
Lessee, Operating Lease, Remaining Lease Term	15 years 7 months 6 days
Maximum [Member] \| Roseville, MN Lease [Member]
Other Commitments [Line Items]
Lessee, Operating Lease, Remaining Lease Term	5 years
Corporate Headquarters [Member]
Other Commitments [Line Items]
Office and lab building area \| ft²	38,000
Equipment [Member]
Other Commitments [Line Items]
Restricted cash \| $	$ 0.2

Leases, Commitments, and Contingencies - Summary of Other Information Related to Leases (Detail) $ in Thousands	9 Months Ended
	Sep. 30, 2022 USD ($)
Cash paid for amounts included in the measurement of lease liabilities:
Operating cash flows, Operating	$ 205
Operating cash flows, Financing	0
Financing cash flows, Operating	0
Financing cash flows, Financing	$ 353
Operating Lease, Weighted average remaining lease term (years)	15 years 7 months 6 days
Finance Lease, Weighted average remaining lease term (years)	7 months 6 days
Operating Lease, Weighted average discount rate	7.90%
Finance Lease, Weighted average discount rate	8.10%

Stock-Based Compensation - Performance Stock Units - Additional Information (Detail) $ in Millions	1 Months Ended	9 Months Ended
	Jul. 31, 2021 Day	Sep. 30, 2022 USD ($) shares	Jul. 07, 2021 shares
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Performance stock, shares granted		2,712,803
2017 Omnibus Plan [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Performance stock, shares granted		3,105,208
Performance Stock Units [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Unrecognized stock-based compensation expense related to performance stock units \| $		$ 1.1
Unrecognized stock-based compensation expense, expected recognition weighted-average period		22 months
Performance Stock Units [Member] \| 2021 Grant \| Inducement Plan [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Vesting period	3 years
Consecutive trading day \| Day	30
Performance Stock Units [Member] \| 2021 Grant \| Mr Carr [Member] \| 2017 Omnibus Plan [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Performance stock, shares granted		530,000
Performance Stock Units [Member] \| 2021 Grant \| Mr Carr [Member] \| Inducement Plan [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Performance stock, shares granted			600,000

Leases, Commitments, and Contingencies - Summary of Non Cancellable Operating Leases (Detail) $ in Thousands	Sep. 30, 2022 USD ($)
Disclosure of Non Cancellable Operating Leases [Line Items]
Right-of-Use Asset	$ 13,734
Roseville, MN Lease [Member]
Disclosure of Non Cancellable Operating Leases [Line Items]
Remaining Term (years)	15 years 7 months 6 days
Right-of-Use Asset	$ 13,734

Related-Party Transactions - Additional Information (Detail) - Cellectis [Member] $ in Millions	9 Months Ended
	Sep. 30, 2022 USD ($)
Related Party Transaction [Line Items]
Minimum net worth required	$ 300
Maximum [Member]
Related Party Transaction [Line Items]
Threshold percentage of ownership in outstanding common stock to enact indemnification agreement	50.00%

Stock-Based Compensation - Summary of Stock-Based Compensation Expenses Related to Restricted Stock Units (Detail) - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021	Sep. 30, 2022	Sep. 30, 2021
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Stock-based compensation expenses	$ 1,035	$ 1,236	$ 2,890	$ 865
Restricted Stock Units [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Stock-based compensation expenses	$ 334	$ 416	$ 992	$ (105)

CONSOLIDATED BALANCE SHEETS (Parenthetical) - $ / shares	Sep. 30, 2022	Dec. 31, 2021
Statement of Financial Position [Abstract]
Common stock, par value	$ 0.0001	$ 0.0001
Common stock, shares authorized	275,000,000	275,000,000
Common stock, shares issued	46,909,420	38,874,146
Common stock, shares outstanding	46,809,268	38,773,994
Treasury stock, shares	100,152	100,152

Stock-Based Compensation (Tables)

9 Months Ended

Sep. 30, 2022

Share-based Compensation Arrangement by Share-based Payment Award [Line Items]

Summary of Fair Values of Stock Options Granted and Assumptions used in Black-Scholes Model

The estimated fair values of stock options granted, and the assumptions used for the Black-Scholes option pricing model were as follows:


	Nine Months Ended September 30,
	2022		2021
Estimated fair values of stock options granted	$	0.86	$	3.93
Assumptions:
Risk-free interest rate		1.9% - 3.5%		0.6% - 1.1%
Expected volatility		89.7% - 92.8%		80.1% - 82.0%
Expected term (in years)		5.50 - 6.89		5.5 - 6.5

Summary of Stock Option Activity

Information on stock option activity is as follows:


	Options Exercisable		Weighted- Average Exercise Price Per Share		Options Outstanding			Weighted- Average Exercise Price Per Share

Balance as of December 31, 2021		2,789,110	$	10.23		4,658,405		$	9.47
Granted						1,609,000			1.12
Exercised						—			—
Forfeited or expired						(420,606	)		7.05

Balance as of September 30, 2022		3,314,828	$	9.88		5,846,799		$	7.35

Summary of Stock-based Compensation Expense

Stock-based compensation expense related to PSUs is as follows:


	Three Months Ended September 30,				Nine Months Ended September 30,
In Thousands	2022		2021		2022		2021

Stock-based compensation expense	$	171	$	122	$	478	$	(133	)

Schedule of Net Cash Proceeds from Exercise of Stock Options Less Shares Used for Minimum Withholding Taxes and Intrinsic Value of Options Exercised

Net cash proceeds from the exercise of stock options less shares used for minimum withholding taxes and the intrinsic value of options exercised were as follows:


	Three Months Ended September 30,				Nine Months Ended September 30,
In Thousands	2022		2021		2022		2021
Net cash proceeds	$	—	$	—	$	—	$	227

Intrinsic value of options exercised	$	—	$	—	$	—	$	344

Summary of Activity of Restricted Stock Units

Information on restricted stock unit activity is as follows:


	Number of Restricted Stock Units Outstanding			Weighted- Average Grant Date Fair Value
Unvested balance as of December 31, 2021		571,303		$	6.15
Granted		1,077,600			1.26
Vested		(274,487	)		6.14
Forfeited		(113,955	)		4.13

Unvested balance as of September 30, 2022		1,260,461		$	2.15

Summary of Grant Date Fair Value of Restricted Stock Unit Awards Vested

The total grant-date fair value of restricted stock unit awards that vested is as follows:


	Three Months Ended September 30,				Nine Months Ended September 30,
In Thousands	2022		2021		2022		2021
Grant-date fair value	$	530	$	581	$	1,686	$	1,223

Summary of Stock-Based Compensation Granted As Deemed Dividends

The Company accounts for stock-based compensation awards granted to employees of Cellectis as deemed dividends. The Company recorded deemed dividends as follows:


	Three Months Ended September 30,					Nine Months Ended September 30,
In Thousands	2022		2021			2022		2021
Deemed dividends from grants to Cellectis employees	$	18	$	(84	)	$	82	$	(55	)

Share-based Payment Arrangement, Performance Shares, Outstanding Activity

PSU activity for the nine months ended September 30, 2022, is as follows:


	Number of PSUs

Outstanding as of December 31, 2021:		745,000
Issued		530,000
Forfeited/canceled		(145,000	)
Exercised		—

Outstanding as of September 30, 2022:		1,130,000

Stock Options [Member]

Share-based Compensation Arrangement by Share-based Payment Award [Line Items]

Summary of Stock-based Compensation Expense

Stock-based compensation expense related to stock option awards is as follows:


	Three Months Ended September 30,				Nine Months Ended September 30,
In Thousands	2022		2021		2022		2021
Stock-based compensation expense	$	530	$	698	$	1,420	$	1,103

Restricted Stock Units [Member]

Share-based Compensation Arrangement by Share-based Payment Award [Line Items]

Summary of Stock-based Compensation Expense

Stock-based compensation expense related to restricted stock units is as follows:


	Three Months Ended September 30,				Nine Months Ended September 30,
In Thousands	2022		2021		2022		2021
Stock-based compensation expense	$	334	$	416	$	992	$	(105	)

Stock-Based Compensation - Summary of Activity of Restricted Stock Units (Detail) - Restricted Stock Units [Member]	9 Months Ended
	Sep. 30, 2022 $ / shares shares
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Number of restricted stock units outstanding, Unvested beginning balance \| shares	571,303
Number of restricted stock units outstanding, Granted \| shares	1,077,600
Number of restricted stock units outstanding, Vested \| shares	(274,487)
Number of restricted stock units outstanding, Forfeited \| shares	(113,955)
Number of restricted stock units outstanding, Unvested ending balance \| shares	1,260,461
Weighted-average grant date fair value, Unvested beginning balance \| $ / shares	$ 6.15
Weighted-average grant date fair value, Granted \| $ / shares	1.26
Weighted-average grant date fair value, Vested \| $ / shares	6.14
Weighted-average grant date fair value, Forfeited \| $ / shares	4.13
Weighted-average grant date fair value, Unvested ending balance \| $ / shares	$ 2.15

Stock-Based Compensation - Summary of Fair Values of Stock Options Granted and Assumptions used in Black-Scholes Model (Detail) - Stock Options [Member] - $ / shares	9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Estimated fair values of stock options granted	$ 0.86	$ 3.93
Minimum [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Risk-free interest rate	1.90%	0.60%
Expected volatility	89.70%	80.10%
Expected term (in years)	5 years 6 months	5 years 6 months
Maximum [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Risk-free interest rate	3.50%	1.10%
Expected volatility	92.80%	82.00%
Expected term (in years)	6 years 10 months 20 days	6 years 6 months

Supplemental Information

9 Months Ended

Sep. 30, 2022

Organization, Consolidation and Presentation of Financial Statements [Abstract]

Supplemental Information

9. SUPPLEMENTAL INFORMATION

Certain statement of operations amounts are as follows:


	Three Months Ended September 30,						Nine Months Ended September 30,
In Thousands	2022			2021			2022			2021
Stock-based compensation expense:
Research and development	$	210		$	252		$	620		$	1,061
Selling, general, and administrative		825			984			2,270			(196	)

Total	$	1,035		$	1,236		$	2,890		$	865


	Three Months Ended September 30,						Nine Months Ended September 30,
In Thousands	2022			2021			2022			2021
Interest, net:
Interest expense	$	(57	)	$	(356	)	$	(73	)	$	(1,075	)
Interest income		31			—			44			16
Common stock warrants - financing costs amortization		(21	)		—			(51	)		—

Total	$	(47	)	$	(356	)	$	(80	)	$	(1,059	)

Certain balance sheet amounts are as follows:


In Thousands	As of September 30, 2022		As of December 31, 2021
Cash, cash equivalents, and restricted cash:
Cash and cash equivalents	$	7,031	$	13,823
Restricted cash		174		499
Non-current restricted cash		—		99

Total	$	7,205	$	14,421

Supplemental statement of cash flows information is as follows:


	As of September 30,
In Thousands	2022		2021
Interest paid	$	67		1,072

Non-cash transactions not reported in the consolidated statement of cash flows is as follows:


	As of September 30,
In Thousands	2022			2021
Receivable from Jefferies for shares issued under ATM Facility	$	(260	)	$	—
Non-cash additions to land, buildings, and equipment	$	(687	)	$	—
Cumulative effect of adoption of lease accounting standard on stockholders’ equity	$	832		$	—
Establishment of operating lease right-of-use assets and associated operating lease liabilities	$	14,090		$	—

CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($) $ in Thousands	9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021
Operating activities
Net loss	$ (14,054)	$ (22,142)
Adjustments to reconcile net loss to net cash used by operating activities:
Gain upon extinguishment of Payroll Protection Program Loan	0	(1,528)
Depreciation and amortization	1,158	1,776
Stock-based compensation	2,890	865
Unrealized (gain) loss on mark-to-market of common stock warrants	(5,009)
Changes in operating assets and liabilities:
Accounts receivable		4,681
Due to/from related parties	(34)	(617)
Inventory		(291)
Prepaid expenses and other current assets	297	2,873
Accounts payable	(188)	108
Accrued expenses	41	(1,742)
Accrued compensation	(166)	334
Other	(536)	1,029
Net cash used by operating activities	(15,601)	(14,654)
Investing activities
Proceeds from sales of short-term investments		11,698
Purchases of land, buildings, and equipment	(1,509)	(376)
Net cash (used by) provided by investing activities	(1,509)	11,322
Financing activities
Proceeds from the issuance of common stock and pre-funded warrants	11,209
Costs incurred related to the issuance of common stock, pre-funded warrants, and common warrants	(962)
Repayments of financing lease obligations	(353)	(271)
Proceeds from the exercise of stock options	0	227
Net cash provided by (used by) financing activities	9,894	(44)
Net (decrease) in cash, cash equivalents, and restricted cash	(7,216)	(3,376)
Cash, cash equivalents and restricted cash - beginning of period	14,421	18,289
Cash, cash equivalents and restricted cash - end of period	$ 7,205	$ 14,913

Leases, Commitments, and Contingencies

9 Months Ended

Sep. 30, 2022

Commitments and Contingencies Disclosure [Abstract]

Leases, Commitments, and Contingencies

8. LEASES, COMMITMENTS, AND CONTINGENCIES

Leases

In February 2016, the FASB issued ASU No. 2016-02, Leases (Topic 842) and in July 2018, ASU No. 2018-10, Codification Improvements to Topic 842, Leases, and ASU 2018-11, Leases (Topic 842) – Targeted Improvements (collectively, the Standard). As discussed in Note 1, the Company adopted the Standard on January 1, 2022.

The Company’s leases are summarized as follows:

•

A lease for its headquarters and laboratory facilities in Roseville, MN which encompasses approximately 38,000 square feet. The original lease term was 20 years, and the Company holds four 5-year renewal options. Historically, this lease was considered a failed sale leaseback based on the nature of the transactions and was reported as a financing-type lease.

•

An equipment financing arrangement that is considered a financing-type lease. This arrangement has a term of four years for each draw. The Company was required to deposit cash into a restricted account in an amount equal to the future rent payments required by the lease. As of September 30, 2022, restricted cash totaled $0.2 million. The Company has the option to request the return of excess collateral annually in December, and the amount the Company expects to receive is reflected as a current asset.

•

A small number of short-term and immaterial leases for office equipment.

The Company’s adoption of the Standard required it to remove its existing land, buildings, and equipment associated with its headquarters lease as well as the associated liability. The Company assessed the elements of its lease agreement and upon adoption, recorded an operating lease associated with the sale leaseback of land underlying the headquarter facility, and a second operating lease associated with the building. The cumulative effect of the adoption of the Standard was recorded to stockholders’ equity. The impact of adoption on the Company’s December 31, 2021, consolidated balance sheet was as follows:


	As Reported December 31, 2021			Adoption of Lease Standard			As Adjusted December 31, 2021
Assets

Land, buildings, and equipment	$	21,731		$	(16,543	)	$	5,188
Operating lease right-of-use assets		—			14,090			14,090

	$	21,731		$	(2,453)		$	19,278

Liabilities and stockholders’ equity
Current portion of financing lease obligations	$	370		$	(4)		$	366
Other current liabilities		191			276			467
Financing lease obligations		17,506			(17,371)			135
Operating lease obligations		—			13,814			13,814
Accumulated deficit		(196,092	)		832			(195,260	)

	$	(178,025	)	$	(2,453	)	$	(180,478	)

The Company records its operating lease liabilities at the present value of the future lease payments over the lease term. If the lease term includes options to extend or terminate the lease, those elements are included in the determination of lease term when it is reasonably certain that the option will be exercised. The rate used to determine the present value of future lease payments is the rate stated in the lease agreement, or if not stated, the Company’s incremental borrowing rate is used, up to an effective rate that enables the lease liability to amortize to zero over the lease term. Rent expense for operating leases is recorded in selling, general, and administrative (SG&A) expense in the consolidated statements of operations and in operating cash flows in the consolidated statements of cash flows. The Company also records operating lease right-of-use assets at an initial amount equal to the operating lease liability. Those right-of-use assets are amortized to lease expense within SG&A over the lease term using the effective interest method to ensure the right-of-use asset amortizes to zero concurrent with the associated liability, and the right-of-use asset amortization expense is also reported in operating cash flows in the consolidated statements of cash flows.

The Company records its financing lease liabilities at the present value of the future lease payments over the lease term. If the lease term includes options to extend or terminate the lease, those elements are included in the determination of lease term when it is reasonably certain that the option will be exercised. The rate used to determine the present value of future lease payments is the rate stated in the lease agreement, or if not stated, the Company’s incremental borrowing rate is used, up to an effective rate that enables the lease liability to amortize to zero over the lease term. Expense associated with financing leases is recorded in interest, net in the consolidated statements of operations and in operating cash flows in the consolidated statements of cash flows.

The Company is obligated under non-cancellable operating leases, primarily for office space and certain equipment, as follows:


	As of September 30, 2022
	Remaining		Right-of-Use
In Thousands	Term (years)		Asset
Roseville, MN lease		15.6	$	13,734
Total			$	13,734

The Roseville, MN lease includes four options to each extend the lease for 5 years. These options to extend the lease are not recognized as part of the right-of-use assets and operating lease liabilities as it is not reasonably certain that the Company will exercise those options. The Company’s agreement does not include options to terminate the lease.

The components of lease expense were as follows:


In Thousands	Three Months Ended September 30, 2022		Nine Months Ended September 30, 2022
Finance lease costs	$	57	$	73
Operating lease costs		381		1,174
Variable lease costs		260		717

Total	$	698	$	1,964

Operating lease cost for short-term leases was not material for the nine months ended September 30, 2022.

Other information related to leases was as follows:


In Thousands except for lease term and discount rate	As of and for Nine Months Ended September 30, 2022
	Operating			Financing
Cash paid for amounts included in the measurement of lease liabilities:

Operating cash flows	$	205		$	—
Financing cash flows	$	—		$	353
Weighted average remaining lease term (years)		15.6			0.6
Weighted average discount rate		7.9	%		8.1	%

As of September 30, 2022, future minimum payments under operating and finance leases were as follows:


In Thousands	Operating			Financing			Total
Remainder of 2022	$	345		$	25		$	370
2023		1,446			100			1,546
2024		1,480			—			1,480
2025		1,479			—			1,479
2026		1,479			—			1,479
2027		1,479			—			1,479
Thereafter		16,991			—			16,991

		24,699			125			24,824
Less: imputed interest		(10,814	)		(5	)		(10,819	)

Total	$	13,885		$	120		$	14,005

Litigation and Claims

The Company is not currently a party to any material pending legal proceeding except as disclosed in Note 10.

Financial instruments and financial risk management - Summary of Fair Value of the Common Warrants (Details)	Sep. 30, 2022 yr
Fair Value Measurement Inputs and Valuation Techniques [Line Items]
Estimated fair value of Common Warrants	0.05
Risk-free interest rate
Fair Value Measurement Inputs and Valuation Techniques [Line Items]
Assumptions	4.1
Expected volatility
Fair Value Measurement Inputs and Valuation Techniques [Line Items]
Assumptions	85
Expected term to liquidation (in years)
Fair Value Measurement Inputs and Valuation Techniques [Line Items]
Assumptions	4.9

Stockholders' Equity - Additional Information (Detail) - USD ($) $ / shares in Units, $ in Millions	Oct. 03, 2022	Feb. 23, 2022	Sep. 30, 2022
Equity, Class of Treasury Stock [Line Items]
Common stock, issued and sold		3,880,000
Net proceeds from issuance of common stock and exercise of overallotment		$ 10.0
After deduction of underwriting discounts and estimated other offering expenses		$ 0.9
Pre-funded Warrants [Member]
Equity, Class of Treasury Stock [Line Items]
Class of Warrant or Right, Exercise Price of Warrants or Rights		$ 0.0001	$ 0
Pre-funded Warrants [Member] \| Maximum [Member]
Equity, Class of Treasury Stock [Line Items]
Warrants issued (in shares)		3,880,000
Common Warrants [Member] \| Maximum [Member]
Equity, Class of Treasury Stock [Line Items]
Warrants issued (in shares)		7,760,000
Common Stock Warrants [Member]
Equity, Class of Treasury Stock [Line Items]
Class of Warrant or Right, Exercise Price of Warrants or Rights		$ 1.41	$ 1.41
Common Stock, Voting Rights		4.99
Common Stock Warrants [Member] \| Maximum [Member]
Equity, Class of Treasury Stock [Line Items]
Date from which Warrants or Rights Exercisable		Aug. 23, 2027
Common Stock Warrants [Member] \| Minimum [Member]
Equity, Class of Treasury Stock [Line Items]
Date from which Warrants or Rights Exercisable		Aug. 23, 2022
Subsequent Event [Member] \| ATM Facility [Member]
Equity, Class of Treasury Stock [Line Items]
Common stock, issued and sold	2,000,000
After deduction of underwriting discounts and estimated other offering expenses	$ 0.1
Subsequent Event [Member] \| ATM Facility [Member] \| Maximum [Member]
Equity, Class of Treasury Stock [Line Items]
Common stock shares subscribed but not issued	15,661,000
Follow-on Public Offering [Member]
Equity, Class of Treasury Stock [Line Items]
Common stock, issued and sold		3,880,000
Common stock issued price per share		$ 1.41
Net proceeds from issuance of common stock and exercise of overallotment		$ 10.0
After deduction of underwriting discounts and estimated other offering expenses		$ 0.9
Follow-on Public Offering [Member] \| Pre-funded Warrants [Member]
Equity, Class of Treasury Stock [Line Items]
Common stock issued price per share		$ 1.4099
Follow-on Public Offering [Member] \| Pre-funded Warrants [Member] \| Maximum [Member]
Equity, Class of Treasury Stock [Line Items]
Warrants issued (in shares)		3,880,000
Follow-on Public Offering [Member] \| Common Warrants [Member] \| Maximum [Member]
Equity, Class of Treasury Stock [Line Items]
Warrants issued (in shares)		7,760,000

Leases, Commitments, and Contingencies - Summary of Lease Cost (Detail) - USD ($) $ in Thousands	3 Months Ended	9 Months Ended
	Sep. 30, 2022	Sep. 30, 2022
Lease, Cost [Abstract]
Finance Lease cost	$ 57	$ 73
Operating lease cost	381	1,174
Variable lease cost	260	717
Total	$ 698	$ 1,964

Supplemental Information - Summary of Certain Statements of Operations Amounts (Detail) - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021	Sep. 30, 2022	Sep. 30, 2021
Share-based Compensation Arrangement by Share-based Payment Award, Compensation Cost [Line Items]
Stock-based compensation expenses	$ 1,035	$ 1,236	$ 2,890	$ 865
Interest expense	(57)	(356)	(73)	(1,075)
Interest income	31	0	44	16
Common stock warrants - financing costs amortization	(21)	0	(51)	0
Total	(47)	(356)	(80)	(1,059)
Research and Development [Member]
Share-based Compensation Arrangement by Share-based Payment Award, Compensation Cost [Line Items]
Stock-based compensation expenses	210	252	620	1,061
Selling, General, and Administrative [Member]
Share-based Compensation Arrangement by Share-based Payment Award, Compensation Cost [Line Items]
Stock-based compensation expenses	$ 825	$ 984	$ 2,270	$ (196)

Supplemental Information - Summary of Certain Balance Sheet Amounts (Detail) - USD ($) $ in Thousands	Dec. 31, 2022	Sep. 30, 2022	Dec. 31, 2021
Condensed Balance Sheet Statements Captions [Line Items]
Cash and cash equivalents		$ 7,031	$ 13,823
Restricted cash	$ 100	174	499
Non-current restricted cash		0	99
Total		$ 7,205	$ 14,421

Stock-Based Compensation - Summary of Stock-Based Compensation Expense Related to Stock Option Awards (Detail) - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021	Sep. 30, 2022	Sep. 30, 2021
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Stock-based compensation expenses	$ 1,035	$ 1,236	$ 2,890	$ 865
Stock Options [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Stock-based compensation expenses	$ 530	$ 698	$ 1,420	$ 1,103

Stock-Based Compensation - Summary of Stock-Based Compensation Expenses Related to Performance Stock Units (Detail) - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021	Sep. 30, 2022	Sep. 30, 2021
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Stock-based compensation expenses	$ 1,035	$ 1,236	$ 2,890	$ 865
Performance Stock Units [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Stock-based compensation expenses	$ 171	$ 122	$ 478	$ (133)

Stock-Based Compensation - Summary of Stock-Based Compensation Granted as Deemed Dividends (Detail) - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021	Sep. 30, 2022	Sep. 30, 2021
Nonemployee Restricted Stock Units [Member] \| Cellectis [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Deemed dividends from grants to Cellectis employee	$ 18	$ (84)	$ 82	$ (55)

Going Concern - Additional Information (Details) - USD ($)			3 Months Ended		9 Months Ended
	Oct. 03, 2022	Feb. 23, 2022	Sep. 30, 2022	Sep. 30, 2021	Sep. 30, 2022	Sep. 30, 2021	Dec. 31, 2022	Dec. 31, 2021	Dec. 31, 2020
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Net loss			$ 5,950,000	$ 7,307,000	$ 14,054,000	$ 22,142,000
Net cash used by operating activities					15,601,000	14,654,000
Cash, cash equivalents, and restricted cash			7,205,000	$ 14,913,000	7,205,000	$ 14,913,000		$ 14,421,000	$ 18,289,000
Cash expense for separation-related payments			174,000		174,000		$ 100,000	499,000
Current liabilities			4,228,000		4,228,000			$ 4,854,000
Common stock, issued and sold		3,880,000
Issuance of common stock		$ 10,000,000
After deduction of underwriting discounts and estimated other offering expenses		$ 900,000
ATM Facility [Member]
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Potential cash inflow from the offer of shares					100,000
Shelf Registration Statement [Member] \| ATM Facility [Member]
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Minimum public float			75,000,000		75,000,000
Pre-funded Warrants [Member] \| Maximum [Member]
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Warrants issued (in shares)		3,880,000
Common Warrants [Member] \| Maximum [Member]
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Warrants issued (in shares)		7,760,000
Subsequent Event [Member] \| ATM Facility [Member]
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Common stock, issued and sold	2,000,000
After deduction of underwriting discounts and estimated other offering expenses	$ 100,000
Subsequent Event [Member] \| Maximum [Member] \| ATM Facility [Member]
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Common stock shares subscribed but not issued	15,661,000
Equipment [Member]
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Restricted cash			$ 200,000		$ 200,000

Stockholders' Equity - Summary Of Warrants Transactions (Detail) - $ / shares	9 Months Ended
	Sep. 30, 2022	Feb. 23, 2022
Pre Funded Warrants [Member]
Disclosure of Warrants Transactions [Line Items]
Number of restricted stock units outstanding, Issued	3,880,000
Weighted-average grant date fair value, Issued	$ 0.0001
Number of restricted stock units outstanding, Exercised	3,880,000
Number of restricted stock units outstanding, Unvested ending balance	0
Weighted-average grant date fair value, Unvested ending balance	$ 0
Class of Warrant or Right, Number of Securities Called by Each Warrant or Right	0
Class of Warrant or Right, Exercise Price of Warrants or Rights	$ 0	$ 0.0001
Common Stock Warrants [Member]
Disclosure of Warrants Transactions [Line Items]
Number of restricted stock units outstanding, Issued	7,760,000
Weighted-average grant date fair value, Issued	$ 1.41
Number of restricted stock units outstanding, Unvested ending balance	7,760,000
Weighted-average grant date fair value, Unvested ending balance	$ 1.41
Class of Warrant or Right, Number of Securities Called by Each Warrant or Right	7,760,000
Class of Warrant or Right, Exercise Price of Warrants or Rights	$ 1.41	$ 1.41

Stockholders' Equity (Tables)

9 Months Ended

Sep. 30, 2022

Text Block [Abstract]

Disclosure of Warrants Transactions

Warrant transactions for the nine months ended September 30, 2022, are as follows:


	Number of Pre-Funded Warrants		Weighted Average Exercise Price		Number of Common Warrants		Weighted Average Exercise Price
Outstanding as of December 31, 2021:

Issued		3,880,000	$	0.0001		7,760,000	$	1.41
Forfeited/canceled		—		—		—		—
Exercised		3,880,000	$	0.0001		—

Outstanding as of September 30, 2022:		—		—		7,760,000	$	1.41

Exercisable as of September 30, 2022:		—		—		7,760,000	$	1.41

Stockholders' Equity

9 Months Ended

Sep. 30, 2022

Federal Home Loan Banks [Abstract]

Stockholders' Equity

5. STOCKHOLDERS’ EQUITY

Follow-On Public Offering

On February 23, 2022, the Company completed the Follow-On Offering, in which it issued 3,880,000 shares of its common stock, Pre-Funded Warrants to purchase up to 3,880,000 shares of its common stock, and Common Warrants to purchase up to 7,760,000 shares of its common stock. The aggregate offering price for each share of common stock and accompanying Common Warrant was $1.41. The aggregate offering price for each Pre-Funded Warrant and accompanying Common Warrant was $1.4099. In the aggregate, the Company received net proceeds of $10.0 million, after deducting approximately $0.9 million of underwriting discounts and estimated other offering expenses.

Pre-Funded Warrants

Each Pre-Funded Warrant entitled the holder to purchase one share of the Company’s common stock at an exercise price of $0.0001 per share. The Pre-Funded Warrants were recorded as a component of stockholders’ equity within additional paid-in capital. The Pre-Funded Warrants were exercised in full on May 4, 2022, and subsequently settled with the counterparty.

Common Stock Warrants

Each Common Warrant entitles the holder to purchase one share of common stock at an exercise price of $1.41 per share. The Common Warrants became exercisable on August 23, 2022 and expire on August 23, 2027. The Common Warrants are recorded as a liability in the Company’s consolidated balance sheet. Per the terms of the Common Warrants, a holder of an outstanding warrant is not entitled to exercise any portion of such warrant if, upon exercise of such portion of the warrant, the holder’s ownership of the Company’s common stock (together with its affiliates) or the combined voting power of the Company’s securities beneficially owned by such holder (together with its affiliates) would exceed the 4.99 percent after giving effect to the exercise.

Warrant transactions for the nine months ended September 30, 2022, are as follows:


	Number of Pre-Funded Warrants		Weighted Average Exercise Price		Number of Common Warrants		Weighted Average Exercise Price
Outstanding as of December 31, 2021:

Issued		3,880,000	$	0.0001		7,760,000	$	1.41
Forfeited/canceled		—		—		—		—
Exercised		3,880,000	$	0.0001		—

Outstanding as of September 30, 2022:		—		—		7,760,000	$	1.41

Exercisable as of September 30, 2022:		—		—		7,760,000	$	1.41

ATM Facility

On September 21, 2021, the Company entered into an ATM Facility with Jefferies LLC, as sole selling agent. On October 3, 2022, the Company entered into an amendment to the Open Market Sale Agreement that enables it, subject to the applicable baby shelf rules, to offer and sell up to 15,661,000

shares of its common stock. At its discretion the Company determines the timing and number of shares to be issued under the ATM Facility. During the nine months ended September 30, 2022, and in the subsequent period through October 3, 2022, the Company did not issue any shares under the ATM Facility. From October 3, 2022, through the date of this report, the Company has issued approximately

2.0

million shares of common stock under the ATM Facility for proceeds of

0.1

million net of commissions and payments for other share issuance costs.

- Definition

The entire disclosure for shareholders' equity comprised of portions attributable to the parent entity and noncontrolling interest, including other comprehensive income. Includes, but is not limited to, balances of common stock, preferred stock, additional paid-in capital, other capital and retained earnings, accumulated balance for each classification of other comprehensive income and amount of comprehensive income.

+ References

Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef
-Publisher FASB
-Name Accounting Standards Codification
-Topic 505
-URI https://asc.fasb.org/topic&trid=2208762

Reference 2: http://www.xbrl.org/2003/role/disclosureRef
-Publisher FASB
-Name Accounting Standards Codification
-Topic 505
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-Section 50
-Paragraph 13
-Subparagraph (b)
-URI https://asc.fasb.org/extlink&oid=126973232&loc=SL123496158-112644

Reference 3: http://www.xbrl.org/2003/role/disclosureRef
-Publisher FASB
-Name Accounting Standards Codification
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-SubTopic 10
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-URI https://asc.fasb.org/extlink&oid=126973232&loc=SL123496158-112644

Reference 4: http://www.xbrl.org/2003/role/disclosureRef
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-Name Accounting Standards Codification
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-Paragraph 14
-Subparagraph (c)
-URI https://asc.fasb.org/extlink&oid=126973232&loc=SL123496171-112644

Reference 5: http://www.xbrl.org/2003/role/disclosureRef
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-Name Accounting Standards Codification
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-Paragraph 14
-Subparagraph (b)
-URI https://asc.fasb.org/extlink&oid=126973232&loc=SL123496171-112644

Reference 6: http://www.xbrl.org/2003/role/disclosureRef
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-Name Accounting Standards Codification
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-Section S99
-Paragraph 1
-Subparagraph (SX 210.3-04)
-URI https://asc.fasb.org/extlink&oid=120397183&loc=d3e187085-122770

Reference 7: http://www.xbrl.org/2003/role/disclosureRef
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-Name Accounting Standards Codification
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-Subparagraph (d)
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Reference 8: http://www.xbrl.org/2003/role/disclosureRef
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Reference 9: http://www.xbrl.org/2003/role/disclosureRef
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-Name Accounting Standards Codification
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Reference 10: http://www.xbrl.org/2003/role/disclosureRef
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Reference 12: http://www.xbrl.org/2003/role/disclosureRef
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Reference 13: http://www.xbrl.org/2003/role/disclosureRef
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-Name Accounting Standards Codification
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-Paragraph 16
-Subparagraph (b)
-URI https://asc.fasb.org/extlink&oid=126973232&loc=SL123496180-112644

Reference 14: http://www.xbrl.org/2003/role/disclosureRef
-Publisher FASB
-Name Accounting Standards Codification
-Topic 505
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-Paragraph 13
-Subparagraph (i)
-URI https://asc.fasb.org/extlink&oid=126973232&loc=SL123496158-112644

Reference 15: http://www.xbrl.org/2003/role/disclosureRef
-Publisher FASB
-Name Accounting Standards Codification
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Reference 16: http://www.xbrl.org/2003/role/disclosureRef
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-Paragraph 6
-Subparagraph (a)
-URI https://asc.fasb.org/extlink&oid=126731327&loc=SL126733271-114008

+ Details

Stock-Based Compensation - Stock Options - Additional Information (Detail) $ in Thousands	9 Months Ended
	Sep. 30, 2022 USD ($)
2017 Omnibus Plan [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Options priced at fair market value, Percent	100.00%
Stock option expiration period	10 years
2017 Omnibus Plan [Member] \| Minimum [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Stock option, vesting period	3 years
2017 Omnibus Plan [Member] \| Maximum [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Stock option, vesting period	6 years
Stock Options [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Weighted average remaining contractual term	5 years 3 months 18 days
Aggregate intrinsic value of options outstanding and exercisable	$ 0
Unrecognized stock-based compensation expense related to non-vested stock options	$ 3,700
Unrecognized stock-based compensation expense, expected recognition weighted-average period	23 months

Basis of Presentation & Summary of Significant Accounting Policies - Additional Information (Detail) - USD ($) $ / shares in Units, $ in Thousands	9 Months Ended
	Sep. 30, 2022	Jun. 30, 2022	Jan. 01, 2022	Dec. 31, 2021	Sep. 30, 2021	Jun. 30, 2021	Dec. 31, 2020
Property, Plant and Equipment [Line Items]
Impairment of long lived assets held for use	$ 0
Operating lease right-of-use assets	13,736			$ 0
Operating lease obligations	13,550			0
Land buildings and equipment and operating lease right of use asset lessee	18,600
Stockholders equity	8,844	$ 13,759		$ 14,132	$ 15,825	$ 21,896	$ 36,875
Market capitalization	8,000
Accounting Standards Update 2016-02 [Member]
Property, Plant and Equipment [Line Items]
Operating lease right-of-use assets			$ 14,100
Operating lease obligations			$ 14,100
Stockholders equity cumulative effect of adoption of lease accounting standard.	$ 800
Follow On Offering [Member]
Property, Plant and Equipment [Line Items]
Class of Warrant or Right, Exercise Price of Warrants or Rights	$ 1.41
Date from which Warrants or Rights Exercisable	Aug. 23, 2027

Financial Instruments Measured at Fair Value and Concentrations of Credit Risk - Summary of Fair Value Measurements and Financial Statement Presentation (Detail) $ in Thousands	Sep. 30, 2022 USD ($)
Fair Value Concentration Of Risk Financial Statement Captions [Line Items]
Fair Values of Liabilities	$ 402
Common Stock Warrants [Member]
Fair Value Concentration Of Risk Financial Statement Captions [Line Items]
Fair Values of Liabilities	402
Level 3 [Member]
Fair Value Concentration Of Risk Financial Statement Captions [Line Items]
Fair Values of Liabilities	402
Level 3 [Member] \| Common Stock Warrants [Member]
Fair Value Concentration Of Risk Financial Statement Captions [Line Items]
Fair Values of Liabilities	$ 402

Subsequent Event - Additional Information (Detail) $ in Thousands	Nov. 30, 2022 USD ($)
Subsequent Event [Line Items]
Litigation settlement, amount awarded from other party	$ 750
Subsequent Event [Member]
Subsequent Event [Line Items]
Litigation settlement, amount awarded from other party	$ 750

Supplemental Information (Tables)

9 Months Ended

Sep. 30, 2022

Organization, Consolidation and Presentation of Financial Statements [Abstract]

Schedule of Certain Statements of Operations Amounts

Certain statement of operations amounts are as follows:


	Three Months Ended September 30,						Nine Months Ended September 30,
In Thousands	2022			2021			2022			2021
Stock-based compensation expense:
Research and development	$	210		$	252		$	620		$	1,061
Selling, general, and administrative		825			984			2,270			(196	)

Total	$	1,035		$	1,236		$	2,890		$	865


	Three Months Ended September 30,						Nine Months Ended September 30,
In Thousands	2022			2021			2022			2021
Interest, net:
Interest expense	$	(57	)	$	(356	)	$	(73	)	$	(1,075	)
Interest income		31			—			44			16
Common stock warrants - financing costs amortization		(21	)		—			(51	)		—

Total	$	(47	)	$	(356	)	$	(80	)	$	(1,059	)

Schedule of Certain Balance Sheet Amounts

Certain balance sheet amounts are as follows:


In Thousands	As of September 30, 2022		As of December 31, 2021
Cash, cash equivalents, and restricted cash:
Cash and cash equivalents	$	7,031	$	13,823
Restricted cash		174		499
Non-current restricted cash		—		99

Total	$	7,205	$	14,421

Schedule of Statements of Certain Statements of Cash Flows Amounts

Supplemental statement of cash flows information is as follows:


	As of September 30,
In Thousands	2022		2021
Interest paid	$	67		1,072

Non-cash transactions not reported in the consolidated statement of cash flows is as follows:


	As of September 30,
In Thousands	2022			2021
Receivable from Jefferies for shares issued under ATM Facility	$	(260	)	$	—
Non-cash additions to land, buildings, and equipment	$	(687	)	$	—
Cumulative effect of adoption of lease accounting standard on stockholders’ equity	$	832		$	—
Establishment of operating lease right-of-use assets and associated operating lease liabilities	$	14,090		$	—

Financial Instruments Measured at Fair Value and Concentrations of Credit Risk (Tables)

9 Months Ended

Sep. 30, 2022

Text Block [Abstract]

Summary of Fair Value Measurements and Financial Statement Presentation

The fair values of the Company’s financial instruments measured at fair value and their respective levels in the fair value hierarchy as of September 30, 2022, were as follows:


	September 30, 2022								September 30, 2022
	Fair Values of Assets								Fair Values of Liabilities
In Thousands	Level 1		Level 2		Level 3		Total		Level 1		Level 2		Level 3		Total
Other items reported at fair value:
Common stock warrants	$	—	$	—	$	—	$	—	$	—	$	—	$	402	$	402

Total	$	—	$	—	$	—	$	—	$	—	$	—	$	402	$	402

Summary of Fair Value of the Common Warrants

The estimated fair values of the Common Warrants, and the assumptions used for the Black-Scholes option pricing model were as follows:


	As of September 30, 2022

Estimated fair value of Common Warrants	$	0.05
Assumptions:
Risk-free interest rate		4.1	%
Expected volatility		85.0	%
Expected term to liquidation (in years)		4.9

Leases, Commitments, and Contingencies - Summary of Future Minimum Rental Payments (Detail) $ in Thousands	Sep. 30, 2022 USD ($)
Schedule of Future Minimum Rental Payments [Abstract]
Operating, Remainder of 2022	$ 345
Operating, 2023	1,446
Operating, 2024	1,480
Operating, 2025	1,479
Operating, 2026	1,479
Operating, 2027	1,479
Operating, Thereafter	16,991
Total Operating	24,699
Operating, Less: imputed interest	(10,814)
Total	13,885
Financing, Remainder of 2022	25
Financing, 2023	100
Financing, 2024	0
Financing, 2025	0
Financing, 2026	0
Financing, 2027	0
Financing, Thereafter	0
Total Financing	125
Financing, Less: imputed interest	(5)
Total	120
Remainder of 2022	370
2023	1,546
2024	1,480
2025	1,479
2026	1,479
2027	1,479
Thereafter	16,991
Total	24,824
Less: imputed interest	(10,819)
Total	$ 14,005

CONSOLIDATED BALANCE SHEETS - USD ($) $ in Thousands	Sep. 30, 2022	Dec. 31, 2021
Current assets:
Cash and cash equivalents	$ 7,031	$ 13,823
Restricted cash	174	499
Prepaid expenses and other current assets	739	859
Total current assets	7,944	15,181
Non-current restricted cash	0	99
Land, buildings, and equipment	4,859	21,731
Operating lease right-of-use assets	13,736	0
Other non-current assets	163	183
Total assets	26,702	37,194
Current liabilities:
Accounts payable	435	1,260
Accrued expenses	380	339
Accrued compensation	2,356	2,522
Due to related parties	138	172
Current portion of financing lease obligations	120	370
Common stock warrants	402	0
Other current liabilities	397	191
Total current liabilities	4,228	4,854
Financing lease obligations	0	17,506
Operating lease obligations	13,550	0
Other non-current liabilities	80	702
Total liabilities	17,858	23,062
Stockholders' equity:
Common stock, $0.0001 par value; 275,000,000 shares authorized; 46,909,420 shares issued and 46,809,268 shares outstanding as of September 30, 2022, and 38,874,146 shares issued and 38,773,994 shares outstanding as of December 31, 2021	5	4
Additional paid-in capital	219,196	211,263
Common stock in treasury, at cost; 100,152 shares as of September 30, 2022, and December 31, 2021	(1,043)	(1,043)
Accumulated deficit	(209,314)	(196,092)
Total stockholders' equity	8,844	14,132
Total liabilities and stockholders' equity	$ 26,702	$ 37,194

Income Taxes	9 Months Ended
Income Taxes	Sep. 30, 2022
Income Tax Disclosure [Abstract]
Income Taxes	7. INCOME TAXES The Company provides for a valuation allowance when it is more likely than not that it will not realize a portion of the deferred tax assets. The Company has established a full valuation allowance for deferred tax assets due to the uncertainty that enough taxable income will be generated in the taxing jurisdiction to utilize the assets. Therefore, the Company has not reflected any benefit of such deferred tax assets in the accompanying consolidated financial statements. As of September 30, 2022, there were no material changes to what the Company disclosed regarding tax uncertainties or penalties as of December 31, 2021.

- Definition

The entire disclosure for income taxes. Disclosures may include net deferred tax liability or asset recognized in an enterprise's statement of financial position, net change during the year in the total valuation allowance, approximate tax effect of each type of temporary difference and carryforward that gives rise to a significant portion of deferred tax liabilities and deferred tax assets, utilization of a tax carryback, and tax uncertainties information.

+ References

Reference 1: http://www.xbrl.org/2003/role/disclosureRef
-Publisher FASB
-Name Accounting Standards Codification
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-Name Accounting Standards Codification
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-Subparagraph (SX 210.4-08(h)(2))
-URI https://asc.fasb.org/extlink&oid=120395691&loc=d3e23780-122690

+ Details

Leases, Commitments, and Contingencies (Tables)

9 Months Ended

Sep. 30, 2022

Lease, Cost [Abstract]

Schedule Of Impact Of Adoption Of The Standard On Consolidated Balance Sheet

The impact of adoption on the Company’s December 31, 2021, consolidated balance sheet was as follows:


	As Reported December 31, 2021			Adoption of Lease Standard			As Adjusted December 31, 2021
Assets

Land, buildings, and equipment	$	21,731		$	(16,543	)	$	5,188
Operating lease right-of-use assets		—			14,090			14,090

	$	21,731		$	(2,453)		$	19,278

Liabilities and stockholders’ equity
Current portion of financing lease obligations	$	370		$	(4)		$	366
Other current liabilities		191			276			467
Financing lease obligations		17,506			(17,371)			135
Operating lease obligations		—			13,814			13,814
Accumulated deficit		(196,092	)		832			(195,260	)

	$	(178,025	)	$	(2,453	)	$	(180,478	)

Summary of Non Cancellable Operating Leases

The Company is obligated under non-cancellable operating leases, primarily for office space and certain equipment, as follows:


	As of September 30, 2022
	Remaining		Right-of-Use
In Thousands	Term (years)		Asset
Roseville, MN lease		15.6	$	13,734
Total			$	13,734

Summary of Lease Cost

The components of lease expense were as follows:


In Thousands	Three Months Ended September 30, 2022		Nine Months Ended September 30, 2022
Finance lease costs	$	57	$	73
Operating lease costs		381		1,174
Variable lease costs		260		717

Total	$	698	$	1,964

Summary of Other Information Related to Leases

Other information related to leases was as follows:


In Thousands except for lease term and discount rate	As of and for Nine Months Ended September 30, 2022
	Operating			Financing
Cash paid for amounts included in the measurement of lease liabilities:

Operating cash flows	$	205		$	—
Financing cash flows	$	—		$	353
Weighted average remaining lease term (years)		15.6			0.6
Weighted average discount rate		7.9	%		8.1	%

Summary of Future Minimum Rental Payments

As of September 30, 2022, future minimum payments under operating and finance leases were as follows:


In Thousands	Operating			Financing			Total
Remainder of 2022	$	345		$	25		$	370
2023		1,446			100			1,546
2024		1,480			—			1,480
2025		1,479			—			1,479
2026		1,479			—			1,479
2027		1,479			—			1,479
Thereafter		16,991			—			16,991

		24,699			125			24,824
Less: imputed interest		(10,814	)		(5	)		(10,819	)

Total	$	13,885		$	120		$	14,005

Supplemental Information - Summary of Statements of Certain Statements of Cash Flows Amounts (Detail) - USD ($) $ in Thousands	9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021
Condensed Financial Information Disclosure [Abstract]
Interest paid	$ 67	$ 1,072
Receivable from Jefferies for shares issued under ATM Facility	(260)	0
Non-cash additions to land, buildings, and equipment	(687)	0
Cumulative effect of adoption of lease accounting standard on stockholders' equity	832	0
Establishment of right-to-use assets and associated operating lease liabilities	$ 14,090	$ 0

Financial Instruments Measured at Fair Value and Concentrations of Credit Risk - Additional Information (Detail) - USD ($) $ in Millions	Sep. 30, 2022	Dec. 31, 2021
Fair Value Concentration Of Risk Financial Statement Captions [Line Items]
Fair value of financing leases		$ 14.5
Corporate Debt Securities [Member]
Fair Value Concentration Of Risk Financial Statement Captions [Line Items]
Other Financial Instruments	$ 0.0

Stock-Based Compensation - Summary of Net Cash Proceeds from Exercise of Stock Options Less Shares Used for Minimum Withholding Taxes and Intrinsic Value of Options Exercised (Detail) - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021	Sep. 30, 2022	Sep. 30, 2021
Share-based Payment Arrangement [Abstract]
Net cash proceeds	$ 0	$ 0	$ 0	$ 227
Intrinsic value of options exercised	$ 0	$ 0	$ 0	$ 344

Cover Page - shares	9 Months Ended
Cover Page - shares	Sep. 30, 2022	Nov. 02, 2022
Document Information [Line Items]
Document Type	10-Q
Amendment Flag	false
Document Quarterly Report	true
Document Period End Date	Sep. 30, 2022
Document Transition Report	false
Document Fiscal Year Focus	2022
Document Fiscal Period Focus	Q3
Entity File Number	001-38161
Entity Registrant Name	Calyxt, Inc.
Entity Central Index Key	0001705843
Entity Incorporation, State or Country Code	DE
Entity Tax Identification Number	27-1967997
Current Fiscal Year End Date	--12-31
Entity Address, Address Line One	2800 Mount Ridge Road
Entity Address, City or Town	Roseville
Entity Address, State or Province	MN
Entity Address, Postal Zip Code	55113-1127
City Area Code	651
Local Phone Number	683-2807
Title of 12(b) Security	Common Stock (0.0001 par value)
Trading Symbol	CLXT
Security Exchange Name	NASDAQ
Entity Current Reporting Status	Yes
Entity Interactive Data Current	Yes
Entity Filer Category	Non-accelerated Filer
Entity Small Business	false
Entity Emerging Growth Company	true
Entity Ex Transition Period	false
Entity Shell Company	false
Entity Common Stock, Shares Outstanding		48,841,286

CONSOLIDATED STATEMENTS OF OPERATIONS - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021	Sep. 30, 2022	Sep. 30, 2021
Income Statement [Abstract]
Revenue	$ 42	$ 7,762	$ 115	$ 24,044
Costs of goods sold	0	8,281	0	26,553
Gross profit	42	(519)	115	(2,509)
Operating expenses:
Research and development	3,016	2,579	9,207	8,473
Selling, general, and administrative	3,229	3,859	9,965	11,640
Total operating expenses	6,245	6,438	19,172	20,113
Loss from operations	(6,203)	(6,957)	(19,057)	(22,622)
Gain upon extinguishment of Payroll Protection Program Loan	0	0	0	1,528
Interest, net	(47)	(356)	(80)	(1,059)
Non-operating income (expenses)	300	6	5,083	11
Loss before income taxes	(5,950)	(7,307)	(14,054)	(22,142)
Income taxes	0	0	0	0
Net loss	$ (5,950)	$ (7,307)	$ (14,054)	$ (22,142)
Earnings Per Share, Basic	$ (0.13)	$ (0.2)	$ (0.31)	$ (0.6)
Earnings Per Share, Diluted	$ (0.13)	$ (0.2)	$ (0.31)	$ (0.6)
Weighted Average Number of Shares Outstanding, Basic	46,784,445	37,279,703	45,173,455	37,205,655
Weighted Average Number of Shares Outstanding, Diluted	46,784,445	37,279,703	45,173,455	37,205,655
Anti-dilutive stock options, restricted stock units, performance stock units, and common stock warrants	15,997,260	5,966,488	15,997,260	5,966,488

Stock-Based Compensation - Restricted Stock Units - Additional Information (Detail) $ in Millions	9 Months Ended
	Sep. 30, 2022 USD ($)
2017 Omnibus Plan [Member] \| Minimum [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Vesting period	3 years
2017 Omnibus Plan [Member] \| Maximum [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Vesting period	6 years
Restricted Stock Units [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Unrecognized stock-based compensation expense related to restricted stock units	$ 1.6
Unrecognized stock-based compensation expense, expected recognition weighted-average period	25 months
Restricted Stock Units [Member] \| 2017 Omnibus Plan [Member] \| Minimum [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Vesting period	3 years
Restricted Stock Units [Member] \| 2017 Omnibus Plan [Member] \| Maximum [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Vesting period	5 years

Financial Instruments Measured at Fair Value and Concentrations of Credit Risk

9 Months Ended

Sep. 30, 2022

Text Block [Abstract]

Financial Instruments Measured at Fair Value and Concentrations of Credit Risk

3. FINANCIAL INSTRUMENTS MEASURED AT FAIR VALUE AND CONCENTRATIONS OF CREDIT RISK

Financial Instruments Measured at Fair Value and Financial Statement Presentation

Financial instruments including cash and cash equivalents, restricted cash, accounts payable, and all other current liabilities have carrying values that approximate fair value. The Company measures common stock warrants on a quarterly basis. The accounting guidance establishes a three-tier hierarchy, which prioritizes the inputs used in the valuation methodologies in measuring fair value as of the measurement date as follows:

Level 1: Fair values are based on unadjusted quoted prices in active trading markets for identical assets and liabilities.

Level 2: Fair values are based on observable quoted prices other than those in Level 1, such as quoted prices for similar assets or liabilities in active markets or quoted prices for identical assets or liabilities in inactive markets.

Level 3: Fair values are based on at least one significant unobservable input for the asset or liability.

Fair Value Measurements and Financial Statement Presentation

The fair values of the Company’s financial instruments measured at fair value and their respective levels in the fair value hierarchy as of September 30, 2022, were as follows:


	September 30, 2022								September 30, 2022
	Fair Values of Assets								Fair Values of Liabilities
In Thousands	Level 1		Level 2		Level 3		Total		Level 1		Level 2		Level 3		Total
Other items reported at fair value:
Common stock warrants	$	—	$	—	$	—	$	—	$	—	$	—	$	402	$	402

Total	$	—	$	—	$	—	$	—	$	—	$	—	$	402	$	402

The Company estimates the fair value of each Common Warrant as of the date of issuance and at the end of every fiscal period using a Black-Scholes option pricing model, which requires it to make predictive assumptions regarding future stock price volatility and dividend yield. The Company estimates the risk-free interest rate based on the United States Treasury zero-coupon yield curve for the remaining life of the Common Warrant. The Company estimates its future stock price volatility using its historical volatility over the remaining life of the Common Warrant. The Company does not pay dividends and does not expect to pay dividends in the foreseeable future.

The estimated fair values of the Common Warrants, and the assumptions used for the Black-Scholes option pricing model were as follows:


	As of September 30, 2022

Estimated fair value of Common Warrants	$	0.05
Assumptions:
Risk-free interest rate		4.1	%
Expected volatility		85.0	%
Expected term to liquidation (in years)		4.9

As of September 30, 2022, the Company had no other financial instruments measured at fair value.

The non-current portion of the Company’s financing lease obligations are also considered a financial instrument, which the Company measures at fair value for disclosure purposes. It is a Level 2 liability and as of December 31, 2021, it had a fair value of

$14.5

million. There is no non-current portion of the financing lease obligation at September 30, 2022.

Foreign Exchange Risk

Foreign currency fluctuations affect the Company’s foreign currency cash flows related primarily to payments to Cellectis. The Company’s principal foreign currency exposure is to the euro. The Company does not hedge these exposures, and it does not believe that the current level of foreign currency risk is significant to its operations.

Concentrations of Credit Risk

The Company invests its cash, cash equivalents, and restricted cash in highly liquid securities and investment funds. The Company diversifies the risk associated with investing in securities by allocating its investments to a diverse portfolio of short-dated, high investment-grade securities, which it classifies as short-term investments that are recorded at fair value in its consolidated financial statements. The Company maintains the credit risk in this portfolio in accordance with its internal policies and if necessary, makes changes to investments to minimize credit risk. The Company has not experienced any counterparty credit losses. As of September 30, 2022, the Company did not hold any short-term investments.

Going Concern	9 Months Ended
Going Concern	Sep. 30, 2022
Organization, Consolidation and Presentation of Financial Statements [Abstract]
Going Concern	2. GOING CONCERN The Company has incurred losses since its inception. The Company’s net loss was $14.1 million for the nine months ended September 30, 2022, and it used $15.6 million of cash for operating activities for the nine months ended September 30, 2022. The Company’s primary sources of liquidity are its cash and cash equivalents, with additional liquidity accessible, subject to market conditions and other factors, including limitations that may apply to the Company under applicable SEC and Nasdaq Capital Market (Nasdaq) regulations, from the capital markets, including under the Open Market Sale Agreement SM with Jefferies LLC (as amended, the ATM Facility). As of September 30, 2022, the Company had $7.2 million of cash, cash equivalents, and restricted cash. The Company’s restricted cash is associated with its equipment financing leases and was $0.2 million as of September 30, 2022, with $0.1 million scheduled to be returned in December 2022. Current liabilities were $4.2 million as of September 30, 2022. On October 3, 2022, the Company entered into an amendment to the Open Market Sale Agreement with Jefferies for the ATM Facility that enables it, subject to the applicable baby shelf rules described below, to offer and sell up to 15,661,000 shares of its common stock. At its discretion, the Company determines the timing and number of shares to be issued under the ATM Facility. During the nine months ended September 30, 2022, and in the subsequent period through October 3, 2022, the Company did not issue any shares under the ATM Facility. From October 3, 2022, through the date of this report, the Company has issued approximately 2.0 million shares of common stock under the ATM Facility for proceeds of $0.1 million net of commissions and payments for other share issuance costs. On February 23, 2022, the Company issued 3,880,000 shares of its common stock, Pre-Funded Warrants to purchase up to 3,880,000 shares of its common stock, and Common Warrants to purchase up to 7,760,000 shares of its common stock in the Follow-On Offering. In the aggregate, the Company received net proceeds of $10.0 million, after deducting approximately $0.9 million of underwriting discounts and estimated other offering expenses. The Company has incurred losses since its inception and anticipates that it will continue to generate losses for the next several years. Over the longer term and until the Company can generate cash flows sufficient to support its operating capital requirements, it expects to finance a portion of future cash needs through (i) cash on hand, (ii) commercialization activities, which may result in various types of revenue streams from (a) future product development agreements and technology licenses, including upfront and milestone payments, annual license fees, and royalties; and (b) product sales from its proprietary BioFactory production system; (iii) government or other third-party funding, (iv) public or private equity or debt financings, (v) the execution of an alternative strategic transaction pursuant to the board of directors’ ongoing evaluation process, or (v) a combination of the foregoing. However, capital generated by commercialization activities, if any, is expected to be received over a period of time and near-term additional capital may not be available on reasonable terms, if at all. Although the Company has access to the ATM Facility, based on the Company’s public float, as of the date of the filing of its Annual Report on Form 10-K for the fiscal year ended December 31, 2021, the Company is only permitted to utilize a “shelf” registration statement for primary offerings, including the registration statement under which the Company’s ATM Facility is operated, subject to Instruction I.B.6 to Form S-3, which is referred to as the “baby shelf” rules. For so long as the Company’s public float is less than $75,000,000, it may not sell more than the equivalent of one-third of its public float during any twelve consecutive months pursuant to the baby shelf rules. While alternative public and private transaction structures may be available, these may require additional time and cost, may result in substantial dilution to existing stockholders, particularly in light of the Company’s current stock price, may impose operational restrictions on the Company, and may not be available on attractive terms or at all. Accordingly, the Company continuously assesses market conditions and available financing alternatives. In addition, on September 22, 2022, the Company announced that the board of directors is evaluating a full range of strategic alternatives to maximize shareholder value, including financing alternatives, merger, reverse merger, other business combinations, sale of assets, licensing, or other transactions. The Company’s ability to continue as a going concern will depend on its ability to obtain additional public or private equity or debt financing, obtain government or private grants and other similar types of funding, or to consummate an alternative strategic transaction, attain further operating efficiencies, reduce or contain expenditures, and, ultimately, to generate revenue. The Company believes that its cash, cash equivalents, and restricted cash as of September 30, 2022, considering the $0.1 million of net proceeds realized from the ATM Facility as well as potential additional proceeds from the ATM Facility (up to the maximum amount permitted under the baby shelf rules), the legal settlement discussed in Note 10 to the consolidated financial statements, and taking into account additional efforts in reassessing its discretionary spending, including the implementation of increasingly stringent cost reduction and other cash focused measures to manage liquidity, is sufficient to fund its operations into the second quarter of 2023. The Company’s management has concluded there is substantial doubt regarding its ability to continue as a going concern because it will need to raise additional capital to support its business plan for a period of 12 months or more from the date of this filing. The accompanying financial statements have been prepared on a going concern basis, which contemplates the realization of assets and satisfaction of liabilities in the ordinary course of business. The financial statements do not include any adjustments relating to the recoverability and classification of recorded asset amounts or the amounts and classification of liabilities that might result from the outcome of the uncertainties described above. Management has implemented various cost reduction and other cash-focused measures to manage liquidity. If the Company is unable to raise additional capital in a sufficient amount or on acceptable terms or to consummate an alternative strategic transaction, the Company may have to implement increasingly stringent cost saving measures and significantly delay, scale back, or cease operations, in part or in full. If the Company raises additional funds through the issuance of additional debt or equity securities, including as part of a strategic alternative, it could result in substantial dilution to its existing stockholders and increased fixed payment obligations, and these securities may have rights senior to those of the Company’s shares of common stock. Any of these events could significantly harm the Company’s business, financial condition, and prospects.

Leases, Commitments, and Contingencies - Schedule Of Impact Of Adoption Of The Standard On Consolidated Balance Sheet (Detail) - USD ($) $ in Thousands	Sep. 30, 2022	Jan. 01, 2022	Dec. 31, 2021
Assets
Land, buildings, and equipment	$ 4,859		$ 21,731
Operating lease right-of-use assets	13,736		0
Liabilities and stockholders' equity
Current portion of financing lease obligations	120		370
Other current liabilities	397		191
Financing lease obligations	0		17,506
Operating lease obligations	13,550		0
Accumulated deficit	$ (209,314)		(196,092)
Accounting Standards Update 2016-02 [Member]
Assets
Operating lease right-of-use assets		$ 14,100
Liabilities and stockholders' equity
Operating lease obligations		$ 14,100
Cumulative Effect, Period of Adoption, Adjusted Balance [Member] \| Accounting Standards Update 2016-02 [Member]
Assets
Land, buildings, and equipment			5,188
Operating lease right-of-use assets			14,090
Total assets			19,278
Liabilities and stockholders' equity
Current portion of financing lease obligations			366
Other current liabilities			467
Financing lease obligations			135
Operating lease obligations			13,814
Accumulated deficit			(195,260)
Total liabilities and stockholders' equity			(180,478)
Previously Reported [Member] \| Accounting Standards Update 2016-02 [Member]
Assets
Land, buildings, and equipment			21,731
Operating lease right-of-use assets			0
Total assets			21,731
Liabilities and stockholders' equity
Current portion of financing lease obligations			370
Other current liabilities			191
Financing lease obligations			17,506
Operating lease obligations			0
Accumulated deficit			(196,092)
Total liabilities and stockholders' equity			(178,025)
Revision of Prior Period, Accounting Standards Update, Adjustment [Member] \| Accounting Standards Update 2016-02 [Member]
Assets
Land, buildings, and equipment			(16,543)
Operating lease right-of-use assets			14,090
Total assets			(2,453)
Liabilities and stockholders' equity
Current portion of financing lease obligations			(4)
Other current liabilities			276
Financing lease obligations			(17,371)
Operating lease obligations			13,814
Accumulated deficit			832
Total liabilities and stockholders' equity			$ (2,453)

Basis of Presentation & Summary of Significant Accounting Policies (Policies)	9 Months Ended
	Sep. 30, 2022
Accounting Policies [Abstract]
Warrants	Warrants The Company issued pre-funded warrants to purchase common stock (Pre-Funded Warrants) in a follow-on offering on February 23, 2022 (the Follow-On Offering). The Pre-Funded Warrants were exercised in full on May 4, 2022, and subsequently settled with the counterparty. The Company also issued Common Warrants in the Follow-On Offering. The Common Warrants expire on August 23, 2027 and are exercisable for one share of the Company’s common stock for $1.41 per share. The Common Warrants have been classified as a liability because they include a put option election available to their holder that is contingently exercisable if the Company enters into a fundamental transaction (Fundamental Transaction), generally described as a “change of control” (the Change of Control Put). If the Change of Control Put is exercised by the holder of a Common Warrant, they may elect to receive either the consideration of the Fundamental Transaction or put the Common Warrant back to the Company in exchange for cash, based on terms and timing specified in the Common Warrant agreement. If the Change of Control Put option is exercised, the Company is required to pay cash to the holder in an amount as determined by the Black Scholes pricing model, with assumptions determined in accordance with the terms of the Common Warrants. The Common Warrants are reported at fair value with changes in fair value reported in earnings. The Company reports the changes in fair value of the Common Warrants in non-operating income (expenses) in its consolidated statements of operations.
Net Loss Per Share	Net Loss Per Share Due to the Company’s net loss position for the three and nine months ended September 30, 2022, and September 30, 2021, all of its outstanding stock options, restricted stock units (RSUs), performance stock units (PSUs), and warrants to purchase common stock (Common Warrants) are considered anti-dilutive and excluded from the calculation of net loss per share. Accordingly, the treasury method was not used in determining the number of anti-dilutive stock options, RSUs, PSUs, or Common Warrants.
Impairment of Long-Lived Assets	Impairment of Long-Lived Assets As of September 30, 2022, the Company had $18.6 million of land, buildings, and equipment and operating lease right-of-use assets and a net book value of $8.8 million. As of September 30, 2022, the Company’s market capitalization was $8.0 million. The Company has a single asset group and reviews its long-lived assets for impairment whenever events or changes in circumstances indicate that the carrying amount of that asset group may not be recoverable. One such impairment indicator is if the Company’s market capitalization is lower than its net book value for an extended period of time. If the impairment tests indicate that the carrying value of the asset group is greater than the expected undiscounted cash flows to be generated by the asset group, further analysis is performed to determine the fair value of the asset group. To the extent the fair value of the asset group is less than its carrying value, an impairment loss is recognized equal to the amount the carrying value exceeds the fair value of the asset group. Fair value is measured using a discounted cash flow model or independent appraisals, as appropriate. As of the date of this filing, management has determined there was no impairment of its long-lived assets based on its financial projections and its stated pursuit of strategic alternatives, which could include various outcomes.
Recently Issued Accounting Pronouncements	Recently Issued Accounting Pronouncements In February 2016, the FASB issued ASU No. 2016-02, Leases (Topic 842) and in July 2018, ASU No. 2018-10, Codification Improvements to Topic 842, Leases, and ASU 2018-11, Leases (Topic 842) – Targeted Improvements (collectively, the Standard). The Standard requires lessees to record assets and liabilities on the balance sheet for all leases with terms longer than 12 months. The Standard establishes a right-of-use model (ROU) that requires a lessee to recognize a ROU asset and lease liability on the balance sheet for all leases with a term longer than 12 months. Leases will be classified as either finance or operating, with the classification affecting the pattern of expense recognition in the income statement. The Company adopted the Standard as of January 1, 2022, using the transition method which does not require revisions to comparative periods. The Company elected to implement the transition package of practical expedients permitted within the Standard, which among other things, allows it to carryforward the historical lease classification. In addition, the Company elected the hindsight practical expedient to determine the lease term for existing leases and it also made an accounting policy election to not record leases with an initial term of 12 months or less on its consolidated balance sheet. The Company’s adoption of the Standard required it to remove the previously reported amounts for land, buildings, and equipment associated with its headquarters and laboratory facility lease as well as the associated liability. The Company assessed the elements of its lease agreement and upon adoption, recorded an operating lease associated with the sale leaseback of land component of the lease, and a second operating lease associated with the building component of the lease. The Company recorded operating lease assets and liabilities of $14.1 million within its consolidated balance sheet as of January 1, 2022. The Standard had no impact on the Company’s consolidated statements of operations or cash flows. The $0.8 million cumulative effect of the adoption of the Standard was recorded to stockholders’ equity. See Note 8 for further information regarding the Company’s leases. In June 2016, the FASB issued ASU No. 2016-13, “Financial Instruments – Credit Losses (Topic 326)” (ASU 2016-13). ASU 2016-13 creates accounting requirements on how to account for credit losses on most financial assets and certain other instruments. This will require the estimation of lifetime expected credit losses and corresponding recognition of allowance for losses on trade and other receivables, loans, and other instruments held at amortized cost. The ASU requires certain recurring disclosures and is effective for annual periods, and interim periods within those annual periods, beginning on or after December 15, 2023. The Company is in the process of analyzing the impact of this standard on its results of operations.

Stock-Based Compensation - Summary of Grant Date Fair Value of Restricted Stock Unit Awards Vested (Detail) - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021	Sep. 30, 2022	Sep. 30, 2021
Restricted Stock Units [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Grant-date fair value	$ 530	$ 581	$ 1,686	$ 1,223

Stock-Based Compensation

9 Months Ended

Sep. 30, 2022

Share-based Payment Arrangement [Abstract]

Stock-Based Compensation

6. STOCK-BASED COMPENSATION

The Company uses broad-based stock plans to attract and retain highly qualified officers and employees and to help ensure that management’s interests are aligned with those of its shareholders. The Company has also granted equity-based awards to directors, nonemployees, and certain employees of Cellectis.

In December 2014, the Company adopted the Calyxt, Inc. Equity Incentive Plan (2014 Plan), which allowed for the grant of stock options, and in June 2017, it adopted the 2017 Omnibus Plan (2017 Plan), which allowed for the grant of stock options, restricted stock units, performance stock units and other types of equity awards. In July 2021, the Company also adopted the Calyxt, Inc. Employee Inducement Incentive Plan (the Inducement Plan), from which PSUs were granted to Michael A. Carr.

On February 19, 2021, James Blome ceased serving as the Company’s Chief Executive Officer. In the nine-month period ended September 30, 2021, the Company recorded a benefit to earnings from a $2.5 million recapture of non-cash stock compensation expense from the forfeiture of Mr. Blome’s unvested stock options, RSUs, and PSUs.

As of September 30, 2022, 2,712,803 shares were registered and available for grant under effective registration statements, while 3,105,208 shares were available for grant in the form of stock options, restricted stock, restricted stock units, and performance stock units under the 2017 Plan. Stock-based awards currently outstanding also include awards granted under the 2014 Plan and the Inducement Plan. No further awards will be granted under either the 2014 Plan or the Inducement Plan.

Stock Options

The estimated fair values of stock options granted, and the assumptions used for the Black-Scholes option pricing model were as follows:


	Nine Months Ended September 30,
	2022		2021
Estimated fair values of stock options granted	$	0.86	$	3.93
Assumptions:
Risk-free interest rate		1.9% - 3.5%		0.6% - 1.1%
Expected volatility		89.7% - 92.8%		80.1% - 82.0%
Expected term (in years)		5.50 - 6.89		5.5 - 6.5

The Company estimates the fair value of each stock option on the grant date, or other measurement date if applicable, using a Black-Scholes option pricing model, which requires it to make predictive assumptions regarding employee exercise behavior, future stock price volatility, and dividend yield. The Company estimates the risk-free interest rate based on the United States Treasury zero-coupon yield curve at the date of grant for the expected term of the option. The Company estimates its future stock price volatility using the weighted-average historical volatility calculated from a group of comparable public companies over the expected term of the option. The expected term of stock options is estimated using the average of the vesting tranches and the contractual life of each grant for employee options, or the simplified method, as the Company has limited historical information to develop reasonable expectations about future exercise patterns and post-vesting employment termination behavior for its stock option grants. The use of the simplified method is dependent upon the type of equity award granted and the term of the award. The Company does not pay dividends and does not expect to pay dividends in the foreseeable future.

Option strike prices are set at 100 percent or more of the closing share price on the date of grant and generally vest over three to six years following the grant date. Options generally expire 10 years after the date of grant.

Information on stock option activity is as follows:


	Options Exercisable		Weighted- Average Exercise Price Per Share		Options Outstanding			Weighted- Average Exercise Price Per Share

Balance as of December 31, 2021		2,789,110	$	10.23		4,658,405		$	9.47
Granted						1,609,000			1.12
Exercised						—			—
Forfeited or expired						(420,606	)		7.05

Balance as of September 30, 2022		3,314,828	$	9.88		5,846,799		$	7.35

Stock-based compensation expense related to stock option awards is as follows:


	Three Months Ended September 30,				Nine Months Ended September 30,
In Thousands	2022		2021		2022		2021
Stock-based compensation expense	$	530	$	698	$	1,420	$	1,103

As of September 30, 2022, options outstanding and exercisable had no aggregate intrinsic value and the weighted average remaining contractual term was 5.3 years as of that date.

Net cash proceeds from the exercise of stock options less shares used for minimum withholding taxes and the intrinsic value of options exercised were as follows:


	Three Months Ended September 30,				Nine Months Ended September 30,
In Thousands	2022		2021		2022		2021
Net cash proceeds	$	—	$	—	$	—	$	227

Intrinsic value of options exercised	$	—	$	—	$	—	$	344

As of September 30, 2022, unrecognized compensation expense related to non-vested stock options was $3.7 million. This expense will be recognized over 23 months on average.

Restricted Stock Units

The Company grants restricted stock units which generally vest over three to five years after the date of grant. Information on restricted stock unit activity is as follows:


	Number of Restricted Stock Units Outstanding			Weighted- Average Grant Date Fair Value
Unvested balance as of December 31, 2021		571,303		$	6.15
Granted		1,077,600			1.26
Vested		(274,487	)		6.14
Forfeited		(113,955	)		4.13

Unvested balance as of September 30, 2022		1,260,461		$	2.15

The total grant-date fair value of restricted stock unit awards that vested is as follows:


	Three Months Ended September 30,				Nine Months Ended September 30,
In Thousands	2022		2021		2022		2021
Grant-date fair value	$	530	$	581	$	1,686	$	1,223

Stock-based compensation expense related to restricted stock units is as follows:


	Three Months Ended September 30,				Nine Months Ended September 30,
In Thousands	2022		2021		2022		2021
Stock-based compensation expense	$	334	$	416	$	992	$	(105	)

As of September 30, 2022, unrecognized compensation expense related to restricted stock units was $1.6 million. This expense will be recognized over 25 months on average.

The Company accounts for stock-based compensation awards granted to employees of Cellectis as deemed dividends. The Company recorded deemed dividends as follows:


	Three Months Ended September 30,					Nine Months Ended September 30,
In Thousands	2022		2021			2022		2021
Deemed dividends from grants to Cellectis employees	$	18	$	(84	)	$	82	$	(55	)

Performance Stock Units

In June 2022, PSU grants made to two executive officers in 2019 were forfeited because the underlying performance criteria were not met. These PSUs contained a market condition and had a five-year service period. The Company will continue to expense these PSUs over the remaining service period.

In March 2022, the Company granted 530,000 PSUs under the 2017 Plan to five employees including four executive officers. The PSUs include three annual performance periods (2022, 2023, and 2024) and target performance levels for each of those periods linked to the achievement of Company objectives as determined annually for the respective period by the Compensation Committee of the Company’s Board of Directors (the Compensation Committee). Once the annual objectives are approved, the associated expense will be recognized on a straight-line basis over the period from the date of grant through the March 15 determination date. Earned awards will be settled in shares of Company stock no later than the March 15 determination date in the following calendar year. The grant date for the tranche of awards linked to 2022 performance is May 4, 2022. Determination of expense for the 2023 and 2024 tranches of PSUs will be made when the associated business objectives are determined.

In July 2021, the Company granted 600,000 PSUs under the Inducement Plan to Mr. Carr. The PSUs will vest if the Company’s stock remains above three specified price levels for thirty calendar days over the three-year performance period. The PSUs will be settled in unrestricted shares of the Company’s common stock on the vesting date.

PSU activity for the nine months ended September 30, 2022, is as follows:


	Number of PSUs

Outstanding as of December 31, 2021:		745,000
Issued		530,000
Forfeited/canceled		(145,000	)
Exercised		—

Outstanding as of September 30, 2022:		1,130,000

Stock-based compensation expense related to PSUs is as follows:


	Three Months Ended September 30,				Nine Months Ended September 30,
In Thousands	2022		2021		2022		2021

Stock-based compensation expense	$	171	$	122	$	478	$	(133	)

As of September 30, 2022, unrecognized compensation expense related to PSUs was $1.1 million. This expense will be recognized over 22 months on average.

Basis of Presentation & Summary of Significant Accounting Policies	9 Months Ended
	Sep. 30, 2022
Accounting Policies [Abstract]
Basis of Presentation & Summary of Significant Accounting Policies	1. BASIS OF PRESENTATION & SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES The unaudited consolidated financial statements of Calyxt, Inc. (Calyxt or the Company) have been prepared in accordance with accounting principles generally accepted in the United States (U.S. GAAP or GAAP) for interim financial information and with the rules and regulations of the Securities and Exchange Commission (SEC) applicable to interim financial statements. In the Company’s opinion, the accompanying consolidated financial statements reflect all adjustments necessary for a fair presentation of its statements of financial position, results of operations, and cash flows for the periods presented but they do not include all of the information and footnotes required by U.S. GAAP for complete financial statements. Except as otherwise disclosed herein, these adjustments consist of normal recurring items. Operating results for interim periods are not necessarily indicative of results that may be expected for the fiscal year as a whole or any other interim period. The preparation of the consolidated financial statements in conformity with U.S. GAAP requires the Company to make estimates and assumptions that affect the reported amounts of assets, liabilities, revenues, expenses, and the related disclosures at the date of the consolidated financial statements and during the reporting period. Actual results could materially differ from these estimates. Certain prior year amounts have been reclassified to conform to the current year presentation. For further information, refer to the audited consolidated financial statements and notes thereto included in the Company’s Annual Report on Form 10-K for the year ended December 31, 2021, filed with the SEC on March 3, 2022. The accompanying Balance Sheet as of December 31, 2021, was derived from the audited consolidated financial statements. This Quarterly Report on Form 10-Q should be read in conjunction with the Company’s consolidated financial statements and notes included in the Annual Report on Form 10-K for the year ended December 31, 2021. Net Loss Per Share Due to the Company’s net loss position for the three and nine months ended September 30, 2022, and September 30, 2021, all of its outstanding stock options, restricted stock units (RSUs), performance stock units (PSUs), and warrants to purchase common stock (Common Warrants) are considered anti-dilutive and excluded from the calculation of net loss per share. Accordingly, the treasury method was not used in determining the number of anti-dilutive stock options, RSUs, PSUs, or Common Warrants. Warrants The Company issued pre-funded warrants to purchase common stock (Pre-Funded Warrants) in a follow-on offering on February 23, 2022 (the Follow-On Offering). The Pre-Funded Warrants were exercised in full on May 4, 2022, and subsequently settled with the counterparty. The Company also issued Common Warrants in the Follow-On Offering. The Common Warrants expire on August 23, 2027 and are exercisable for one share of the Company’s common stock for $1.41 per share. The Common Warrants have been classified as a liability because they include a put option election available to their holder that is contingently exercisable if the Company enters into a fundamental transaction (Fundamental Transaction), generally described as a “change of control” (the Change of Control Put). If the Change of Control Put is exercised by the holder of a Common Warrant, they may elect to receive either the consideration of the Fundamental Transaction or put the Common Warrant back to the Company in exchange for cash, based on terms and timing specified in the Common Warrant agreement. If the Change of Control Put option is exercised, the Company is required to pay cash to the holder in an amount as determined by the Black Scholes pricing model, with assumptions determined in accordance with the terms of the Common Warrants. The Common Warrants are reported at fair value with changes in fair value reported in earnings. The Company reports the changes in fair value of the Common Warrants in non-operating income (expenses) in its consolidated statements of operations. Impairment of Long-Lived Assets As of September 30, 2022, the Company had $18.6 million of land, buildings, and equipment and operating lease right-of-use assets and a net book value of $8.8 million. As of September 30, 2022, the Company’s market capitalization was $8.0 million. The Company has a single asset group and reviews its long-lived assets for impairment whenever events or changes in circumstances indicate that the carrying amount of that asset group may not be recoverable. One such impairment indicator is if the Company’s market capitalization is lower than its net book value for an extended period of time. If the impairment tests indicate that the carrying value of the asset group is greater than the expected undiscounted cash flows to be generated by the asset group, further analysis is performed to determine the fair value of the asset group. To the extent the fair value of the asset group is less than its carrying value, an impairment loss is recognized equal to the amount the carrying value exceeds the fair value of the asset group. Fair value is measured using a discounted cash flow model or independent appraisals, as appropriate. As of the date of this filing, management has determined there was no impairment of its long-lived assets based on its financial projections and its stated pursuit of strategic alternatives, which could include various outcomes. Recently Issued Accounting Pronouncements In February 2016, the FASB issued ASU No. 2016-02, Leases (Topic 842) and in July 2018, ASU No. 2018-10, Codification Improvements to Topic 842, Leases, and ASU 2018-11, Leases (Topic 842) – Targeted Improvements (collectively, the Standard). The Standard requires lessees to record assets and liabilities on the balance sheet for all leases with terms longer than 12 months. The Standard establishes a right-of-use model (ROU) that requires a lessee to recognize a ROU asset and lease liability on the balance sheet for all leases with a term longer than 12 months. Leases will be classified as either finance or operating, with the classification affecting the pattern of expense recognition in the income statement. The Company adopted the Standard as of January 1, 2022, using the transition method which does not require revisions to comparative periods. The Company elected to implement the transition package of practical expedients permitted within the Standard, which among other things, allows it to carryforward the historical lease classification. In addition, the Company elected the hindsight practical expedient to determine the lease term for existing leases and it also made an accounting policy election to not record leases with an initial term of 12 months or less on its consolidated balance sheet. The Company’s adoption of the Standard required it to remove the previously reported amounts for land, buildings, and equipment associated with its headquarters and laboratory facility lease as well as the associated liability. The Company assessed the elements of its lease agreement and upon adoption, recorded an operating lease associated with the sale leaseback of land component of the lease, and a second operating lease associated with the building component of the lease. The Company recorded operating lease assets and liabilities of $14.1 million within its consolidated balance sheet as of January 1, 2022. The Standard had no impact on the Company’s consolidated statements of operations or cash flows. The $0.8 million cumulative effect of the adoption of the Standard was recorded to stockholders’ equity. See Note 8 for further information regarding the Company’s leases. In June 2016, the FASB issued ASU No. 2016-13, “Financial Instruments – Credit Losses (Topic 326)” (ASU 2016-13). ASU 2016-13 creates accounting requirements on how to account for credit losses on most financial assets and certain other instruments. This will require the estimation of lifetime expected credit losses and corresponding recognition of allowance for losses on trade and other receivables, loans, and other instruments held at amortized cost. The ASU requires certain recurring disclosures and is effective for annual periods, and interim periods within those annual periods, beginning on or after December 15, 2023. The Company is in the process of analyzing the impact of this standard on its results of operations.

CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY (Parenthetical) $ in Millions	9 Months Ended
	Sep. 30, 2022 USD ($)
Follow-on Public Offering [Member]
Adjustments to Additional Paid In Capital Stock And Warrants Issued Issuance Costs	$ 0.5

Related-Party Transactions	9 Months Ended
Related-Party Transactions	Sep. 30, 2022
Related Party Transactions [Abstract]
Related-Party Transactions	4. RELATED-PARTY TRANSACTIONS The Company is party to several agreements that govern its relationship with Cellectis, some of which require the Company to make payments to Cellectis. Pursuant to the Company’s management services agreement with Cellectis, it incurred no management fee expenses for the three and nine months ended September 30, 2022, and it incurred nominal management fee expenses for the three and nine months ended September 30, 2021. Cellectis has also guaranteed the lease agreement for the Company’s headquarters. Cellectis’ guarantee of the Company’s obligations under the lease will terminate at the end of the second consecutive calendar year in which the Company’s tangible net worth exceeds $300 million. The Company agreed to indemnify Cellectis for any obligations incurred by Cellectis under its guaranty of the obligations under the lease, effective upon Cellectis’ ownership falling to 50 percent or less of the Company’s outstanding common stock. This indemnification obligation was triggered in October 2022. TALEN ® is the Company’s primary gene editing technology. TALEN ® technology was invented by researchers at the University of Minnesota and Iowa State University and exclusively licensed to Cellectis. The Company obtained an exclusive license for the TALEN ® technology for commercial use in plants from Cellectis. The Company also licenses other technology from Cellectis. Cellectis is entitled to royalties on any revenue the Company generates from sales of products less certain amounts as defined in the license agreement, royalties on certain cumulative revenue thresholds, and a percentage of any sublicense revenues. The Company has incurred nominal license and royalty fees for the three and nine months ended September 30, 2022 and 2021.

Subsequent Event	9 Months Ended
Subsequent Event	Sep. 30, 2022
Subsequent Events [Abstract]
Subsequent Event	10. SUBSEQUENT EVENT In early November 2022, the Company reached a settlement with one of its technology vendors regarding alleged intellectual property infringement. As a result of the settlement, the Company will receive $0.75 million upon execution of an amended master services agreement (MSA) and another $0.75 million by January 31, 2023. The execution of the MSA is anticipated to occur in late 2022.

Stock-Based Compensation - Additional Information (Detail) - USD ($) $ in Millions	Sep. 30, 2022	Feb. 19, 2021
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Share-based compensation arrangement by share-based payment award, number of shares registered and available for grant	2,712,803
Mr. Blome [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Expected benefit to earnings from recapture of non-cash stock compensation expense		$ 2.5
2017 Omnibus Plan [Member]
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]
Share-based compensation arrangement by share-based payment award, number of shares registered and available for grant	3,105,208

CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($) $ in Thousands	Total	ATM Facility Member	Share-based Payment Arrangement [Member]	IPO [Member]	Common Stock [Member]	Common Stock [Member] IPO [Member]	Additional Paid-In Capital [Member]	Additional Paid-In Capital [Member] ATM Facility Member	Additional Paid-In Capital [Member] Share-based Payment Arrangement [Member]	Additional Paid-In Capital [Member] IPO [Member]	Shares in Treasury [Member]	Accumulated Deficit [Member]
Beginning balance at Dec. 31, 2020	$ 36,875				$ 4		$ 204,807				$ (1,043)	$ (166,893)
Beginning balance, shares at Dec. 31, 2020					37,065,044
Net loss	(22,142)											(22,142)
Stock-based compensation	865						865
Issuance of common stock and payment of minimum employee taxes withheld upon net share settlement of restricted stock units, shares			236,680
Issuance of common stock and payment of minimum employee taxes withheld upon net share settlement of restricted stock units, amount			$ 227						$ 227
Ending balance at Sep. 30, 2021	15,825				$ 4		205,899				(1,043)	(189,035)
Ending balance, shares at Sep. 30, 2021					37,301,724
Beginning balance at Jun. 30, 2021	21,896				$ 4		204,663				(1,043)	(181,728)
Beginning balance, shares at Jun. 30, 2021					37,205,473
Net loss	(7,307)											(7,307)
Stock-based compensation	1,236						1,236
Issuance of common stock and payment of minimum employee taxes withheld upon net share settlement of restricted stock units, shares			96,251
Issuance of common stock and payment of minimum employee taxes withheld upon net share settlement of restricted stock units, amount			$ 0						0
Ending balance at Sep. 30, 2021	15,825				$ 4		205,899				(1,043)	(189,035)
Ending balance, shares at Sep. 30, 2021					37,301,724
Beginning balance at Dec. 31, 2021	14,132				$ 4		211,263				(1,043)	(196,092)
Beginning balance, shares at Dec. 31, 2021					38,773,994
Net loss	(14,054)											(14,054)
Stock-based compensation	2,890						2,890
Issuance of common stock		$ (7)		$ 5,051		$ 1		$ (7)		$ 5,050
Issuance of common stock, shares				3,880,000
Issuance of common stock upon exercise of pre-funded warrants				3,880,000
Issuance of common stock and payment of minimum employee taxes withheld upon net share settlement of restricted stock units, shares			275,274
Cumulative Effect Of Adoption Of Lease Accounting Standard	832											832
Ending balance at Sep. 30, 2022	8,844				$ 5		219,196				(1,043)	(209,314)
Ending balance, shares at Sep. 30, 2022					46,809,268
Beginning balance at Jun. 30, 2022	13,759				$ 5		218,161				(1,043)	(203,364)
Beginning balance, shares at Jun. 30, 2022					46,715,542
Net loss	(5,950)											(5,950)
Stock-based compensation	1,035						1,035
Issuance of common stock and payment of minimum employee taxes withheld upon net share settlement of restricted stock units, shares			93,726
Issuance of common stock and payment of minimum employee taxes withheld upon net share settlement of restricted stock units, amount			$ 0						$ 0
Ending balance at Sep. 30, 2022	$ 8,844				$ 5		$ 219,196				$ (1,043)	$ (209,314)
Ending balance, shares at Sep. 30, 2022					46,809,268

Stock-Based Compensation - Summary of Stock Option Activity (Detail)	9 Months Ended
	Sep. 30, 2022 $ / shares shares
Share-based Payment Arrangement [Abstract]
Options Exercisable, Beginning Balance \| shares	2,789,110
Options Exercisable, Ending Balance \| shares	3,314,828
Weighted-Average Exercise Price Per Share, Beginning Balance \| $ / shares	$ 10.23
Weighted-Average Exercise Price Per Share, Ending Balance \| $ / shares	$ 9.88
Options Outstanding, Beginning Balance \| shares	4,658,405
Options Outstanding, Granted \| shares	1,609,000
Options Outstanding, Forfeited or expired \| shares	(420,606)
Options Outstanding, Ending Balance \| shares	5,846,799
Weighted-Average Exercise Price Per Share, Beginning Balance \| $ / shares	$ 9.47
Weighted-Average Exercise Price Per Share, Granted \| $ / shares	1.12
Weighted-Average Exercise Price Per Share, Forfeited or expired \| $ / shares	7.05
Weighted-Average Exercise Price Per Share, Ending Balance \| $ / shares	$ 7.35